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The modern evolutionary synthesis (often referred to simply as the new synthesis, the modern synthesis, the evolutionary synthesis, neo-Darwinian synthesis or neo-Darwinism), generally denotes the integration of Charles Darwin's theory of the evolution of species by natural selection, Gregor Mendel's theory of genetics as the basis for biological inheritance, random genetic mutation as the source of variation, and mathematical population genetics. Major figures in the development of the modern synthesis include Thomas Hunt Morgan, R. A. Fisher, Theodosius Dobzhansky, J.B.S. Haldane, Sewall Wright, Julian Huxley, Ernst Mayr, Bernhard Rensch, George Gaylord Simpson, and G. Ledyard Stebbins.